if the valve directly below the pumps is open it could be that either pump running will draw through both pipes, the pipe on the left is the hot water tank, if the pump on the riight is the return from the heating then it will be drawing through that too
Under normal circumstances I would expect the tap between the two pumps to be closed. If one pump failed that tsp could be opened to share the pumping between water and heating circuit.
The tsp on the floor serves no purpose as there is another further up the pipe doing the same job.